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Lophocolea bicuspidata

Steph. (1922)

This name is accepted

Kingdom: Viridiplantae Phylum: Marchantiophyta Class/Clade: Jungermanniopsida Order: Jungermanniales Family: Lophocoleaceae Genus: Lophocolea

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Description

Key Characters:

Growth Form:

Leaves/Thallus:

Stems:

Sporophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Indigenous

Island Status

Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Bibliography

Name Published In: Sp. Hepat. (Stephani) 6: 263 (1922)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Lophocolea bicuspidata West Maui, Wst Maui Natural Area Reserve - Panaewa Section, headwaters of Halona Stream Aspect: W. Opportunistic collecting along gulch in mesic forest. Mestic forest: Metrosideros polymorpha, Cheirodendron trigynum, Myrsine lessertiana, Ilex anomala, Xylosma hawaiiense, Hydrangea arguta, Clermontia kakeana, Cyanea elliptica, Cibotium glaucum, Sadleria cyatheoides, Touchardia oahuensis, Touchardia latifolia, Pipturus albidus, Boehmeria grandis, Crytandra munroi, Cyanea scabra, Freycinetia arborea, Neraudia seicea, Nothocestrum longifollum, Diplazium sandwichianum, Dryopteris fuscoatra, Ctentis latifrons, Microlepia strigosa, Dicranopteris linearis Preserved_Specimen Pezzillo, Z. 418 Maui BISH 2023-04-14
2 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6401ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1192 Maui BISH 2025-05-21
3 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. On twigs in spray of waterfall. Preserved_Specimen With Leptoscyphus baldwinii, Riccardia, Megaceros flagellaris (spores green. Gemmae on margins and upper surface of thallus. Freire, A.V. 2023129 Maui BISH 2023-02-23
4 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 597 Maui BISH 2024-03-06
5 Lophocolea bicuspidata Along Mt Kaala Rd, uphill from the water tanks and downhill from where the Dupont Trail meets Mt Kaala Rd, M.K. Thomas collections on this date, 09 NOV 2022 mostly from S side of road (some collections from N side). Steep wet rock-cut face ranging from 1-9 m tall below overhanging patches of Machaerina angustifolia. With Dryopteris fusco-atra, Cibotium, Begonia foliosa, Trematolobelia kaalae, Plantago, Metrosideros polymorpha, Pipturus albidus, Gunnera, Pilea peploides, introduced Adiantum, Peperomia cookiana, Hydrangea arguta. Preserved_Specimen Thomas, M.K. 530 Oahu BISH 2022-11-09
6 Lophocolea bicuspidata Waikamoi Preserve On dead wood, side of gulch. Preserved_Specimen Hall, J.B. 78 Maui BISH 1994-09-04
7 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge. Nauhi Gulch. Ca. 1 km mauka of Nauhi Cabin. Deep moist gulch shaded by remnant Metrosideros Polymorpha forest. Preserved_Specimen Freire, A.V. 2023310 Hawaii BISH 2023-03-25
8 Lophocolea bicuspidata On Terence Spencer - Gayle Chavez tract on northeast side of Wright Road. Forest of Metrosideros polymorpha and Cibotium glaucum Preserved_Specimen Freire, A.V. 2023482 Hawaii BISH 2023-08-16
9 Lophocolea bicuspidata National Park, Kipuka Puaulu, Bird Park Preserved_Specimen Svihla, R.D. 2044 Hawaii BISH 1956-01-06
10 Lophocolea bicuspidata End of Wright Road, below entrance to National Park, Hilo side Preserved_Specimen Also determined as Cryptolophocolea ciliolata, Lejeunea, and Plagiochila Svihla, R.D. 2034 Hawaii BISH 1956-01-06
11 Lophocolea bicuspidata Palakea fence line Preserved_Specimen plus moss Forbes, C.N. 1011 Hawaii BISH 1915-06-29
12 Lophocolea bicuspidata near Thurston Lava Tube Preserved_Specimen under leaves 3-4x as wide as stem Cooray, R.G. 71011412 Hawaii BISH 1971-01-14
13 Lophocolea bicuspidata Kokee, behind cabin Maile Preserved_Specimen cf. hawaiicus Svihla, R.D. 2073 Kauai BISH 1956-02-11
14 Lophocolea bicuspidata Kokee, behind cabin Maile Preserved_Specimen cf. hawaiicus Svihla, R.D. 2070 Kauai BISH 1956-02-11
15 Lophocolea bicuspidata Koolau Mts. Puu Konahuanui Trail Preserved_Specimen Lophocolea fat bidentate lvs; no trigones w. lvs 4-fid, large, connected on only 1 side Bishop, L.E. 127044 Oahu BISH 1970-12-05
16 Lophocolea bicuspidata Kokee, road to YWCA camp Preserved_Specimen Perianth 3-angled Svihla, R.D. 2065 Kauai BISH 1956-02-10
17 Lophocolea bicuspidata Waianae Mts., Palehua Preserved_Specimen Cooke Jr., C.M. 8 Oahu BISH 1898-07-01
18 Lophocolea bicuspidata End of Wright Road, below entrance to National Park, Hilo side Preserved_Specimen Svihla, R.D. 2011 Hawaii BISH 1956-01-06
19 Lophocolea bicuspidata near Thurston Lava Tube Preserved_Specimen Cooray, R.G. 71011401 Hawaii BISH 1971-01-14
20 Lophocolea bicuspidata Settlement Trail Preserved_Specimen Forbes, C.N. 678 Molokai BISH 1915-03-29
21 Lophocolea bicuspidata near Thurston Lava Tube Preserved_Specimen Cooray, R.G. 71011328 Hawaii BISH 1971-01-13
22 Lophocolea bicuspidata Mt. Kaala Preserved_Specimen W/ moss (Thuidium(?)) Svihla, R.D. 3456 Oahu BISH 1934-06-28
23 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Road On bark. Forest of Metrosideros polymorpha and Cibotium glaucum Preserved_Specimen Freire, A.V. 2022599 Hawaii BISH 2022-06-16
24 Lophocolea bicuspidata Kokee, behind cabin Maile Preserved_Specimen cf. hawaiicus Svihla, R.D. 2074 Kauai BISH 1956-02-11
25 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge. Nauhi Gulch, Ca.1 km mauka of Nauhi Cabin. Deep moist gulch shaded by remnant Metrosideros Polymorpha forest. Preserved_Specimen Freire, A.V. 2023291 Hawaii BISH 2023-03-25
26 Lophocolea bicuspidata Kupaua. Ahupuaa of Waikiki. Southeastern Koolau Mountains. Epiphytic on shrubs and on ground in mesic/wet forest. Growing with Freycinetia arborea, Metrosideros polymorpha, Citharexylum caudatum, Kadua affinis, Dicranopteris linearis, and others. Preserved_Specimen Thomas, M.K. 768 Oahu BISH 2024-09-19
27 Lophocolea bicuspidata road to Haleakala Preserved_Specimen Svihla, R.D. 1741 Maui BISH 1955-12-30
28 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 598 Maui BISH 2024-03-06
29 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Road Forest of Metrosideros polymorpha and Cibotium glaucum. Preserved_Specimen Freire, A.V. 2022612 Hawaii BISH 2022-06-16
30 Lophocolea bicuspidata near Thurston Lava Tube Preserved_Specimen Cooray, R.G. 71011404 Hawaii BISH 1971-01-14
31 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6403ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1194 Maui BISH 2025-05-21
32 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1495 Maui BISH 1919-07-01
33 Lophocolea bicuspidata Halemanu Preserved_Specimen Cooke Jr., C.M. 84 Kauai BISH 1903-10-24
34 Lophocolea bicuspidata Palakea fence line Preserved_Specimen Forbes, C.N. 1012 Hawaii BISH 1915-06-29
35 Lophocolea bicuspidata Kaala Preserved_Specimen Svihla, R.D. 3419 Oahu BISH 1934-06-28
36 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1490 Maui BISH 1919-07-01
37 Lophocolea bicuspidata Mt. Kaala Preserved_Specimen Svihla, R.D. 34447 Oahu BISH 1934-06-28
38 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1485 Maui BISH 1919-07-01
39 Lophocolea bicuspidata Keaku Preserved_Specimen Cooke Jr., C.M. 54 Kauai BISH 1903-10-25
40 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1474 Maui BISH 1919-07-01
41 Lophocolea bicuspidata Kaala Preserved_Specimen Svihla, R.D. 3471 Oahu BISH 1934-06-28
42 Lophocolea bicuspidata Waimea Drainage Basin Preserved_Specimen Forbes, C.N. 1581 Kauai BISH 1917-07-03
43 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1484 Maui BISH 1919-07-01
44 Lophocolea bicuspidata Kokee Camp. Trail to Halemanu Preserved_Specimen Svihla, R.D. 2104 Kauai BISH 1956-02-19
45 Lophocolea bicuspidata National Park, Kipuka Puaulu, Bird Park Preserved_Specimen Svihla, R.D. 2047 Hawaii BISH 1956-01-06
46 Lophocolea bicuspidata Waianae Mountain Range: Waianae Kai Preserved_Specimen Higashino, P.K. 331 Oahu BISH 1975-03-15
47 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge. Honohima Gulch, Refuge Headquarters Area. Deep moist gulch shaded by remnant forest. Preserved_Specimen Commonest leaft liverwort in the Refuge. Freire, A.V. 2023202 Hawaii BISH 2023-03-24
48 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6400ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1191 Maui BISH 2025-05-21
49 Lophocolea bicuspidata Lower Waiakamoi rainforest Preserved_Specimen Svihla, R.D. 1892 Maui BISH 1956-01-03
50 Lophocolea bicuspidata below Kokee, trail Preserved_Specimen Svihla, R.D. 2108 Kauai BISH 1956-02-21
51 Lophocolea bicuspidata Palakea fence line Preserved_Specimen Forbes, C.N. 1007 Hawaii BISH 1915-06-29
52 Lophocolea bicuspidata Olinda Preserved_Specimen Svihla, R.D. 34149 Maui BISH 1934-07-19
53 Lophocolea bicuspidata Kokee camps. Road to YWCA camp Preserved_Specimen Svihla, R.D. 2063 Kauai BISH 1956-02-10
54 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge. Nauhi Gulch, Ca.1 km mauka of Nauhi Cabin. Deep moist gulch shaded by remnant Metrosideros Polymorpha forest. Preserved_Specimen Freire, A.V. 2023290 Hawaii BISH 2023-03-25
55 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. Preserved_Specimen Freire, A.V. 202367 Maui BISH 2023-02-23
56 Lophocolea bicuspidata East Maui, The Nature Conservancy of Hawai'I's Waikamoi Preserve, Ainahou bowl at the top of Ko'olau Gap, 5600, Aspect: N. General collections from soiuthern pali of Ainahou bowl. Wet: Metrosideros, Cheirodendron, Leptecophylla, Melicope, Myrsine, Ilex, Hydrangea, Rubus, Coprosma, Pipturus, Smilax, Hoiokula, Diplazium, Dryopteris, Pteris, Athyrium, Polystichum, Astella, Coniogramme, Lobelia Preserved_Specimen Pezzillo, Z. 952 Maui BISH 2024-08-29
57 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge, in and near Awehi Gulch, Metrosideros polymorpha forest below Puu Akala cabin, including deep moist forested gulch. Preserved_Specimen A. Lophocolea bicuspidata Steph.; B. Plagiochila. Leaves toothed.; C. Odontoschisma denudatum (Mart.) Dumort. Freire, A.V. 2023241 Hawaii BISH 2023-03-24
58 Lophocolea bicuspidata Olaa Forest Tract; 1 km NE of the first bend in Wright Road Montane rain forest. On Cibotium, fairly dense shade. Preserved_Specimen Succubous, underleaves deeply 4-lobed, attached to substrate. Burton, P.J. 510 Hawaii BISH 1979-04-08
59 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6402ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1193 Maui BISH 2025-05-21
60 Lophocolea bicuspidata Puu Makaala State Reserve Forest of Metrosideros polymorpha and Cibotium glaucum Preserved_Specimen Leaves quadrate, underleaves poorly developed Freire, A.V. 2019112.5 Hawaii BISH 2019-10-31
61 Lophocolea bicuspidata Ahupuaa of Honouliuli, ili aina of Popouwela. Central Waianae Mountains. Area also known as Kaluaa. Growing in mesic forest dominated by Toona ciliata, Psidium cattleianum, and Schinus terebinthifolius. Also growing with Ceodes umbellifera, C. Brunoniana, Planchonella sandwicensis, Cryptocaria oahuensis, Microlepia setosa, Asplenium macraei, Miconia crenata, Rubus rosifolius, and others. Preserved_Specimen Thomas, M.K. 865 Oahu BISH 2024-10-30
62 Lophocolea bicuspidata Manuka State Wayside, on Highway 11 about Mile 82. Loop trail from parking lot clockwise to forested pit crater. 19° 06'32-7'05 N, 155° 48'50-49'33 W. Disturbed native Metrosideros forest with Ochna serrulata common in understory Preserved_Specimen Freire, A.V. 2022852 Hawaii BISH 2022-09-29
63 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge, in and near 'Awehi Gulch. Forest below Pu'u 'Akala cabin. Metrosideros polymorpha forest, including deep moist forested gulch. Preserved_Specimen Freire, A.V. 2023268 Hawaii BISH
64 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Rd Forest of Metrosideros polymorpha and Cibotium glaucum. Preserved_Specimen With perianths Freire, A.V. 2022169 Hawaii BISH 2022-01-25
65 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge, in and near Awehi Gulch; below Puu Akala cabin Metrosideros polymorpha forest below Puu Akala cabin, including deep moist forested gulch Preserved_Specimen No trigones. Freire, A.V. 2023383 Hawaii BISH 2023-03-26
66 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. Preserved_Specimen With unusual moss and Leptoscyphus baldwinii. Freire, A.V. 2023115 Maui BISH 2023-02-23
67 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Rd Forest of Metrosideros polymorpha and Cibotium glaucum. Preserved_Specimen but small underleaves Freire, A.V. 2022176 Hawaii BISH 2022-01-25
68 Lophocolea bicuspidata On Terence Spencer - Gayle Chavez tract on northeast side of Wright Road. Forest of Metrosideros polymorpha and Cibotium glaucum Preserved_Specimen Freire, A.V. 2023479 Hawaii BISH 2023-08-16
69 Lophocolea bicuspidata S Kohala Mixture on trunks of Cibotium chamissoi. Preserved_Specimen Lohman, M.L. 526 Hawaii BISH 1952-08-28
70 Lophocolea bicuspidata West Maui, Hana'ula, Forest above windmills, in drainages east and west of trail leading up to springs, 0, Aspect: S. Wet mesic forest with DicLin ridges and diverse ferny gulches, speciments opportunistically collected from ground, boulders, and trunks of native species. Preserved_Specimen Pezzillo, Z. 811 Maui BISH 2024-03-15
71 Lophocolea bicuspidata South corner of 10th and Jade Streets, Volcano On Metrosideros polymorpha Preserved_Specimen Freire, A.V. 20201133 Hawaii BISH 2020-12-15
72 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6408ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1199 Maui BISH 2025-05-21
73 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge, in and near Awehi Gulch, Metrosideros polymorpha forest below Puu Akala cabin, including deep moist forested gulch. Preserved_Specimen With numerous perianths Freire, A.V. 2023254 Hawaii BISH 2023-03-24
74 Lophocolea bicuspidata On forest trail Mauna Kapu north to summit of Palikea, Waianae Mts. Locally common on Metrosideros bark on forest trail. Exotic trees dominant for most of trail, grading into Metosideros polymorpha forest at summit. Preserved_Specimen Judziewicz, E.J. 21965 Oahu BISH 2021-11-04
75 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 583 Maui BISH 2024-03-06
76 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 590 Maui BISH 2024-03-06
77 Lophocolea bicuspidata Hawaiian Islands, Maui, West Maui, Waihe'e Valley. Lower shelf near back of valley on south side Aspect: N. Low elevation wet forest, collections found on ground and on wet trunks of assorted native species, heavily invaded by MicCre. Wet Forest: Metrosidros, Cheirodendron, Kadua, Clermontia, Coprosma, Miconia crenata, Diplazium, Asplenium, Paspalum, Freycinetia, Elaphoglossum, Cyanea, Hydrangea, Cyrtandra, Dicranopteris, Rubus, Microsorum, Sadleria, Agertina, Hoiokula, Antidesma, Leisorus, Cibotium, Stenogyne, Athyrium Preserved_Specimen Pezzillo, Z. 701 Maui BISH 2024-03-12
78 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. Preserved_Specimen In moss matrix. Freire, A.V. 202363 Maui BISH 2023-02-23
79 Lophocolea bicuspidata Summit Mt. Kaala On soil Preserved_Specimen Iwatsuki, Z. 22070 Oahu BISH 1974-06-20
80 Lophocolea bicuspidata On forest trail from Mauna Kapu north to summit of Palikea, Waianae Mts. On Metrosideros bark on forest trail. Exotic trees dominant for most of trail, grading into Metosideros polymorpha forest at summit. With Odontoschisma denudata, Cuspidatula robusta. Preserved_Specimen Judziewicz, E.J. 21970 Oahu BISH 2021-11-04
81 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. On twigs in spray of waterfall. Preserved_Specimen Freire, A.V. 2023159 Maui BISH 2023-02-23
82 Lophocolea bicuspidata Puu Makaala Natural Area Reserve, border of Puu Unit of HAVO In tiny Carex alligata bog Preserved_Specimen Freire, A.V. 2022908 Hawaii BISH 2022-10-25
83 Lophocolea bicuspidata West Maui, Hana'ula, Forest above windmills, in drainages east and west of trail leading up to springs, 0, Aspect: S. Wet mesic forest with DicLin ridges and diverse ferny gulches, speciments opportunistically collected from ground, boulders, and trunks of native species. Preserved_Specimen Pezzillo, Z. 804 Maui BISH 2024-03-15
84 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 579 Maui BISH 2024-03-06
85 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Rd Forest of Metrosideros polymorpha and Cibotium glaucum. On Hymenophyllum lanceolatum, with other leafies. Preserved_Specimen Freire, A.V. 2022167 Hawaii BISH 2022-01-25
86 Lophocolea bicuspidata West Maui, Wst Maui Natural Area Reserve - Panaewa Section, headwaters of Halona Stream Aspect: W. Opportunistic collecting along gulch in mesic forest. Mestic forest: Metrosideros polymorpha, Cheirodendron trigynum, Myrsine lessertiana, Ilex anomala, Xylosma hawaiiense, Hydrangea arguta, Clermontia kakeana, Cyanea elliptica, Cibotium glaucum, Sadleria cyatheoides, Touchardia oahuensis, Touchardia latifolia, Pipturus albidus, Boehmeria grandis, Crytandra munroi, Cyanea scabra, Freycinetia arborea, Neraudia seicea, Nothocestrum longifollum, Diplazium sandwichianum, Dryopteris fuscoatra, Ctentis latifrons, Microlepia strigosa, Dicranopteris linearis Preserved_Specimen Pezzillo, Z. 418 Maui BISH 2023-04-14
87 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6401ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1192 Maui BISH 2025-05-21
88 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. On twigs in spray of waterfall. Preserved_Specimen With Leptoscyphus baldwinii, Riccardia, Megaceros flagellaris (spores green. Gemmae on margins and upper surface of thallus. Freire, A.V. 2023129 Maui BISH 2023-02-23
89 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 597 Maui BISH 2024-03-06
90 Lophocolea bicuspidata Along Mt Kaala Rd, uphill from the water tanks and downhill from where the Dupont Trail meets Mt Kaala Rd, M.K. Thomas collections on this date, 09 NOV 2022 mostly from S side of road (some collections from N side). Steep wet rock-cut face ranging from 1-9 m tall below overhanging patches of Machaerina angustifolia. With Dryopteris fusco-atra, Cibotium, Begonia foliosa, Trematolobelia kaalae, Plantago, Metrosideros polymorpha, Pipturus albidus, Gunnera, Pilea peploides, introduced Adiantum, Peperomia cookiana, Hydrangea arguta. Preserved_Specimen Thomas, M.K. 530 Oahu BISH 2022-11-09
91 Lophocolea bicuspidata Waikamoi Preserve On dead wood, side of gulch. Preserved_Specimen Hall, J.B. 78 Maui BISH 1994-09-04
92 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge. Nauhi Gulch. Ca. 1 km mauka of Nauhi Cabin. Deep moist gulch shaded by remnant Metrosideros Polymorpha forest. Preserved_Specimen Freire, A.V. 2023310 Hawaii BISH 2023-03-25
93 Lophocolea bicuspidata On Terence Spencer - Gayle Chavez tract on northeast side of Wright Road. Forest of Metrosideros polymorpha and Cibotium glaucum Preserved_Specimen Freire, A.V. 2023482 Hawaii BISH 2023-08-16
94 Lophocolea bicuspidata National Park, Kipuka Puaulu, Bird Park Preserved_Specimen Svihla, R.D. 2044 Hawaii BISH 1956-01-06
95 Lophocolea bicuspidata End of Wright Road, below entrance to National Park, Hilo side Preserved_Specimen Also determined as Cryptolophocolea ciliolata, Lejeunea, and Plagiochila Svihla, R.D. 2034 Hawaii BISH 1956-01-06
96 Lophocolea bicuspidata Palakea fence line Preserved_Specimen plus moss Forbes, C.N. 1011 Hawaii BISH 1915-06-29
97 Lophocolea bicuspidata near Thurston Lava Tube Preserved_Specimen under leaves 3-4x as wide as stem Cooray, R.G. 71011412 Hawaii BISH 1971-01-14
98 Lophocolea bicuspidata Kokee, behind cabin Maile Preserved_Specimen cf. hawaiicus Svihla, R.D. 2073 Kauai BISH 1956-02-11
99 Lophocolea bicuspidata Kokee, behind cabin Maile Preserved_Specimen cf. hawaiicus Svihla, R.D. 2070 Kauai BISH 1956-02-11
100 Lophocolea bicuspidata Koolau Mts. Puu Konahuanui Trail Preserved_Specimen Lophocolea fat bidentate lvs; no trigones w. lvs 4-fid, large, connected on only 1 side Bishop, L.E. 127044 Oahu BISH 1970-12-05
101 Lophocolea bicuspidata Kokee, road to YWCA camp Preserved_Specimen Perianth 3-angled Svihla, R.D. 2065 Kauai BISH 1956-02-10
102 Lophocolea bicuspidata Waianae Mts., Palehua Preserved_Specimen Cooke Jr., C.M. 8 Oahu BISH 1898-07-01
103 Lophocolea bicuspidata End of Wright Road, below entrance to National Park, Hilo side Preserved_Specimen Svihla, R.D. 2011 Hawaii BISH 1956-01-06
104 Lophocolea bicuspidata near Thurston Lava Tube Preserved_Specimen Cooray, R.G. 71011401 Hawaii BISH 1971-01-14
105 Lophocolea bicuspidata Settlement Trail Preserved_Specimen Forbes, C.N. 678 Molokai BISH 1915-03-29
106 Lophocolea bicuspidata near Thurston Lava Tube Preserved_Specimen Cooray, R.G. 71011328 Hawaii BISH 1971-01-13
107 Lophocolea bicuspidata Mt. Kaala Preserved_Specimen W/ moss (Thuidium(?)) Svihla, R.D. 3456 Oahu BISH 1934-06-28
108 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Road On bark. Forest of Metrosideros polymorpha and Cibotium glaucum Preserved_Specimen Freire, A.V. 2022599 Hawaii BISH 2022-06-16
109 Lophocolea bicuspidata Kokee, behind cabin Maile Preserved_Specimen cf. hawaiicus Svihla, R.D. 2074 Kauai BISH 1956-02-11
110 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge. Nauhi Gulch, Ca.1 km mauka of Nauhi Cabin. Deep moist gulch shaded by remnant Metrosideros Polymorpha forest. Preserved_Specimen Freire, A.V. 2023291 Hawaii BISH 2023-03-25
111 Lophocolea bicuspidata Kupaua. Ahupuaa of Waikiki. Southeastern Koolau Mountains. Epiphytic on shrubs and on ground in mesic/wet forest. Growing with Freycinetia arborea, Metrosideros polymorpha, Citharexylum caudatum, Kadua affinis, Dicranopteris linearis, and others. Preserved_Specimen Thomas, M.K. 768 Oahu BISH 2024-09-19
112 Lophocolea bicuspidata road to Haleakala Preserved_Specimen Svihla, R.D. 1741 Maui BISH 1955-12-30
113 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 598 Maui BISH 2024-03-06
114 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Road Forest of Metrosideros polymorpha and Cibotium glaucum. Preserved_Specimen Freire, A.V. 2022612 Hawaii BISH 2022-06-16
115 Lophocolea bicuspidata near Thurston Lava Tube Preserved_Specimen Cooray, R.G. 71011404 Hawaii BISH 1971-01-14
116 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6403ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1194 Maui BISH 2025-05-21
117 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1495 Maui BISH 1919-07-01
118 Lophocolea bicuspidata Halemanu Preserved_Specimen Cooke Jr., C.M. 84 Kauai BISH 1903-10-24
119 Lophocolea bicuspidata Palakea fence line Preserved_Specimen Forbes, C.N. 1012 Hawaii BISH 1915-06-29
120 Lophocolea bicuspidata Kaala Preserved_Specimen Svihla, R.D. 3419 Oahu BISH 1934-06-28
121 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1490 Maui BISH 1919-07-01
122 Lophocolea bicuspidata Mt. Kaala Preserved_Specimen Svihla, R.D. 34447 Oahu BISH 1934-06-28
123 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1485 Maui BISH 1919-07-01
124 Lophocolea bicuspidata Keaku Preserved_Specimen Cooke Jr., C.M. 54 Kauai BISH 1903-10-25
125 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1474 Maui BISH 1919-07-01
126 Lophocolea bicuspidata Kaala Preserved_Specimen Svihla, R.D. 3471 Oahu BISH 1934-06-28
127 Lophocolea bicuspidata Waimea Drainage Basin Preserved_Specimen Forbes, C.N. 1581 Kauai BISH 1917-07-03
128 Lophocolea bicuspidata Ukulele Preserved_Specimen Forbes, C.N. 1484 Maui BISH 1919-07-01
129 Lophocolea bicuspidata Kokee Camp. Trail to Halemanu Preserved_Specimen Svihla, R.D. 2104 Kauai BISH 1956-02-19
130 Lophocolea bicuspidata National Park, Kipuka Puaulu, Bird Park Preserved_Specimen Svihla, R.D. 2047 Hawaii BISH 1956-01-06
131 Lophocolea bicuspidata Waianae Mountain Range: Waianae Kai Preserved_Specimen Higashino, P.K. 331 Oahu BISH 1975-03-15
132 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge. Honohima Gulch, Refuge Headquarters Area. Deep moist gulch shaded by remnant forest. Preserved_Specimen Commonest leaft liverwort in the Refuge. Freire, A.V. 2023202 Hawaii BISH 2023-03-24
133 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6400ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1191 Maui BISH 2025-05-21
134 Lophocolea bicuspidata Lower Waiakamoi rainforest Preserved_Specimen Svihla, R.D. 1892 Maui BISH 1956-01-03
135 Lophocolea bicuspidata below Kokee, trail Preserved_Specimen Svihla, R.D. 2108 Kauai BISH 1956-02-21
136 Lophocolea bicuspidata Palakea fence line Preserved_Specimen Forbes, C.N. 1007 Hawaii BISH 1915-06-29
137 Lophocolea bicuspidata Olinda Preserved_Specimen Svihla, R.D. 34149 Maui BISH 1934-07-19
138 Lophocolea bicuspidata Kokee camps. Road to YWCA camp Preserved_Specimen Svihla, R.D. 2063 Kauai BISH 1956-02-10
139 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge. Nauhi Gulch, Ca.1 km mauka of Nauhi Cabin. Deep moist gulch shaded by remnant Metrosideros Polymorpha forest. Preserved_Specimen Freire, A.V. 2023290 Hawaii BISH 2023-03-25
140 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. Preserved_Specimen Freire, A.V. 202367 Maui BISH 2023-02-23
141 Lophocolea bicuspidata East Maui, The Nature Conservancy of Hawai'I's Waikamoi Preserve, Ainahou bowl at the top of Ko'olau Gap, 5600, Aspect: N. General collections from soiuthern pali of Ainahou bowl. Wet: Metrosideros, Cheirodendron, Leptecophylla, Melicope, Myrsine, Ilex, Hydrangea, Rubus, Coprosma, Pipturus, Smilax, Hoiokula, Diplazium, Dryopteris, Pteris, Athyrium, Polystichum, Astella, Coniogramme, Lobelia Preserved_Specimen Pezzillo, Z. 952 Maui BISH 2024-08-29
142 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge, in and near Awehi Gulch, Metrosideros polymorpha forest below Puu Akala cabin, including deep moist forested gulch. Preserved_Specimen A. Lophocolea bicuspidata Steph.; B. Plagiochila. Leaves toothed.; C. Odontoschisma denudatum (Mart.) Dumort. Freire, A.V. 2023241 Hawaii BISH 2023-03-24
143 Lophocolea bicuspidata Olaa Forest Tract; 1 km NE of the first bend in Wright Road Montane rain forest. On Cibotium, fairly dense shade. Preserved_Specimen Succubous, underleaves deeply 4-lobed, attached to substrate. Burton, P.J. 510 Hawaii BISH 1979-04-08
144 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6402ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1193 Maui BISH 2025-05-21
145 Lophocolea bicuspidata Puu Makaala State Reserve Forest of Metrosideros polymorpha and Cibotium glaucum Preserved_Specimen Leaves quadrate, underleaves poorly developed Freire, A.V. 2019112.5 Hawaii BISH 2019-10-31
146 Lophocolea bicuspidata Ahupuaa of Honouliuli, ili aina of Popouwela. Central Waianae Mountains. Area also known as Kaluaa. Growing in mesic forest dominated by Toona ciliata, Psidium cattleianum, and Schinus terebinthifolius. Also growing with Ceodes umbellifera, C. Brunoniana, Planchonella sandwicensis, Cryptocaria oahuensis, Microlepia setosa, Asplenium macraei, Miconia crenata, Rubus rosifolius, and others. Preserved_Specimen Thomas, M.K. 865 Oahu BISH 2024-10-30
147 Lophocolea bicuspidata Manuka State Wayside, on Highway 11 about Mile 82. Loop trail from parking lot clockwise to forested pit crater. 19° 06'32-7'05 N, 155° 48'50-49'33 W. Disturbed native Metrosideros forest with Ochna serrulata common in understory Preserved_Specimen Freire, A.V. 2022852 Hawaii BISH 2022-09-29
148 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge, in and near 'Awehi Gulch. Forest below Pu'u 'Akala cabin. Metrosideros polymorpha forest, including deep moist forested gulch. Preserved_Specimen Freire, A.V. 2023268 Hawaii BISH
149 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Rd Forest of Metrosideros polymorpha and Cibotium glaucum. Preserved_Specimen With perianths Freire, A.V. 2022169 Hawaii BISH 2022-01-25
150 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge, in and near Awehi Gulch; below Puu Akala cabin Metrosideros polymorpha forest below Puu Akala cabin, including deep moist forested gulch Preserved_Specimen No trigones. Freire, A.V. 2023383 Hawaii BISH 2023-03-26
151 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. Preserved_Specimen With unusual moss and Leptoscyphus baldwinii. Freire, A.V. 2023115 Maui BISH 2023-02-23
152 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Rd Forest of Metrosideros polymorpha and Cibotium glaucum. Preserved_Specimen but small underleaves Freire, A.V. 2022176 Hawaii BISH 2022-01-25
153 Lophocolea bicuspidata On Terence Spencer - Gayle Chavez tract on northeast side of Wright Road. Forest of Metrosideros polymorpha and Cibotium glaucum Preserved_Specimen Freire, A.V. 2023479 Hawaii BISH 2023-08-16
154 Lophocolea bicuspidata S Kohala Mixture on trunks of Cibotium chamissoi. Preserved_Specimen Lohman, M.L. 526 Hawaii BISH 1952-08-28
155 Lophocolea bicuspidata West Maui, Hana'ula, Forest above windmills, in drainages east and west of trail leading up to springs, 0, Aspect: S. Wet mesic forest with DicLin ridges and diverse ferny gulches, speciments opportunistically collected from ground, boulders, and trunks of native species. Preserved_Specimen Pezzillo, Z. 811 Maui BISH 2024-03-15
156 Lophocolea bicuspidata South corner of 10th and Jade Streets, Volcano On Metrosideros polymorpha Preserved_Specimen Freire, A.V. 20201133 Hawaii BISH 2020-12-15
157 Lophocolea bicuspidata USA, Hawaiian Islands, Maui, East Maui, TNC's Waikamoi Preserve, Upper Waikamoi, west of Honomanu gulch along contour trail, edge of forest and shrubland, 6408ft, Aspect: NE. Opportunistic collections in subalpine shrubland. Subalpine shrubland: Vaccinium reticulatum, Oreobolus, Lycopodium, Metrosideros, Carex alligat, Carex uncinata, Dryopteris glabra hobdyana, Leptecophylla, Pteridium, Sadleria cyatheoides, Coprosma ernodeoides, Coprosma montana. Preserved_Specimen Pezzillo, Z. 1199 Maui BISH 2025-05-21
158 Lophocolea bicuspidata Hakalau Forest National Wildlife Refuge, in and near Awehi Gulch, Metrosideros polymorpha forest below Puu Akala cabin, including deep moist forested gulch. Preserved_Specimen With numerous perianths Freire, A.V. 2023254 Hawaii BISH 2023-03-24
159 Lophocolea bicuspidata On forest trail Mauna Kapu north to summit of Palikea, Waianae Mts. Locally common on Metrosideros bark on forest trail. Exotic trees dominant for most of trail, grading into Metosideros polymorpha forest at summit. Preserved_Specimen Judziewicz, E.J. 21965 Oahu BISH 2021-11-04
160 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 583 Maui BISH 2024-03-06
161 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 590 Maui BISH 2024-03-06
162 Lophocolea bicuspidata Hawaiian Islands, Maui, West Maui, Waihe'e Valley. Lower shelf near back of valley on south side Aspect: N. Low elevation wet forest, collections found on ground and on wet trunks of assorted native species, heavily invaded by MicCre. Wet Forest: Metrosidros, Cheirodendron, Kadua, Clermontia, Coprosma, Miconia crenata, Diplazium, Asplenium, Paspalum, Freycinetia, Elaphoglossum, Cyanea, Hydrangea, Cyrtandra, Dicranopteris, Rubus, Microsorum, Sadleria, Agertina, Hoiokula, Antidesma, Leisorus, Cibotium, Stenogyne, Athyrium Preserved_Specimen Pezzillo, Z. 701 Maui BISH 2024-03-12
163 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. Preserved_Specimen In moss matrix. Freire, A.V. 202363 Maui BISH 2023-02-23
164 Lophocolea bicuspidata Summit Mt. Kaala On soil Preserved_Specimen Iwatsuki, Z. 22070 Oahu BISH 1974-06-20
165 Lophocolea bicuspidata On forest trail from Mauna Kapu north to summit of Palikea, Waianae Mts. On Metrosideros bark on forest trail. Exotic trees dominant for most of trail, grading into Metosideros polymorpha forest at summit. With Odontoschisma denudata, Cuspidatula robusta. Preserved_Specimen Judziewicz, E.J. 21970 Oahu BISH 2021-11-04
166 Lophocolea bicuspidata TNC Waikamoi Preserve, western fork of Honomanu Stream. Wet montane forest of planted conifers (upper elevations) to native Metrosideros-dominated forest at lower elevations. On twigs in spray of waterfall. Preserved_Specimen Freire, A.V. 2023159 Maui BISH 2023-02-23
167 Lophocolea bicuspidata Puu Makaala Natural Area Reserve, border of Puu Unit of HAVO In tiny Carex alligata bog Preserved_Specimen Freire, A.V. 2022908 Hawaii BISH 2022-10-25
168 Lophocolea bicuspidata West Maui, Hana'ula, Forest above windmills, in drainages east and west of trail leading up to springs, 0, Aspect: S. Wet mesic forest with DicLin ridges and diverse ferny gulches, speciments opportunistically collected from ground, boulders, and trunks of native species. Preserved_Specimen Pezzillo, Z. 804 Maui BISH 2024-03-15
169 Lophocolea bicuspidata East Maui, Kula Forest Reserve, between Plum Trail and lower boundry Collections made in very remnant mesic forest that has mostly been filled in by introduced forestry plantings and introduced understory species Preserved_Specimen Pezzillo, Z. 579 Maui BISH 2024-03-06
170 Lophocolea bicuspidata Puu Makaala State Reserve NW of end of Amaumau Rd Forest of Metrosideros polymorpha and Cibotium glaucum. On Hymenophyllum lanceolatum, with other leafies. Preserved_Specimen Freire, A.V. 2022167 Hawaii BISH 2022-01-25
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